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The One, also known as the Prime Program, is a systemic feature of the Matrix wherein a special code is carried by a randomly selected human being. This person is then gifted vast superhuman abilities as an avatar within the Matrix, which sometimes manifests as extraordinary powers in the real world. The special code has the function to collectively attach, to the person that carries it, all of the anomalies which emanate from the few humans who continue to reject the Matrix. This essentially makes The One an "Integral Anomaly" or the SUM of all anomalies.
The mechanisms by which The One appears are rather simple and arranged in the Prime Program process: when the anomalies in the Matrix reach a certain threshold and begin to pose serious problems, a random human is selected by the machines to be born with a special code (the Integral Anomaly) that, as said before, ties and attaches all of the anomalies within the Matrix to the programming of this human. The Oracle, who is aware of The One's existence and purpose because it is her duty to guide humanity to find The One, will enlighten him to his true nature and lead him towards the Source where he fulfills his purpose of reinserting the Prime Program and resetting the Matrix. She does this by predicting the return of The One to the people of Zion to aid them in freeing humanity from the Machines' grip. The Architect, as creator of the Matrix, is also well aware of The One's existence and purpose, referring to The One as an "eventuality".
Essentially, the purpose of The One is to ensure the stability of the Matrix through containing the anomalies in one individual rather than letting them spread throughout the system.
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